More Letters of Charles Darwin — Volume 2. Darwin Charles
of land has been successively covered with a beech forest between the Caucasus and Japan!
I have not yet seen (for I have not sent to the station) Falconer's works. When you say that you sigh to think how poor your reprinted memoirs would appear, on my soul I should like to shake you till your bones rattled for talking such nonsense. Do you sigh over the "Insular Floras," the Introduction to New Zealand Flora, to Australia, your Arctic Flora, and dear Galapagos, etc., etc., etc.? In imagination I am grinding my teeth and choking you till I put sense into you. Farewell. I have amused myself by writing an audaciously long letter. By the way, we heard yesterday that George has won the second Smith's Prize, which I am excessively glad of, as the Second Wrangler by no means always succeeds. The examination consists exclusively of {the} most difficult subjects, which such men as Stokes, Cayley, and Adams can set.
LETTER 385. A.R. WALLACE TO CHARLES DARWIN. March 8th, 1868.
...While writing a few pages on the northern alpine forms of plants on the Java mountains I wanted a few cases to refer to like Teneriffe, where there are no northern forms and scarcely any alpine. I expected the volcanoes of Hawaii would be a good case, and asked Dr. Seemann about them. It seems a man has lately published a list of Hawaiian plants, and the mountains swarm with European alpine genera and some species! (385/1. "This turns out to be inaccurate, or greatly exaggerated. There are no true alpines, and the European genera are comparatively few. See my 'Island Life,' page 323." — A.R.W.) Is not this most extraordinary, and a puzzler? They are, I believe, truly oceanic islands, in the absence of mammals and the extreme poverty of birds and insects, and they are within the Tropics.
Will not that be a hard nut for you when you come to treat in detail on geographical distribution? I enclose Seemann's note, which please return when you have copied the list, if of any use to you.
LETTER 386. TO J.D. HOOKER. Down, February 21st {1870}.
I read yesterday the notes on Round Island (386/1. In Wallace's "Island Life," page 410, Round Island is described as an islet "only about a mile across, and situated about fourteen miles north-east of Mauritius." Wallace mentions a snake, a python belonging to the peculiar and distinct genus Casarea, as found on Round Island, and nowhere else in the world. The palm Latania Loddigesii is quoted by Wallace as "confined to Round Island and two other adjacent islets." See Baker's "Flora of the Mauritius and the Seychelles." Mr. Wallace says that, judging from the soundings, Round Island was connected with Mauritius, and that when it was "first separated {it} would have been both much larger and much nearer the main island.") which I owe to you. Was there ever such an enigma? If, in the course of a week or two, you can find time to let me hear what you think, I should very much like to hear: or we hope to be at Erasmus' on March 4th for a week. Would there be any chance of your coming to luncheon then? What a case it is. Palms, screw-pines, four snakes — not one being in main island — lizards, insects, and not one land bird. But, above everything, such a proportion of individual monocotyledons! The conditions do not seem very different from the Tuff Galapagos Island, but, as far as I remember, very few monocotyledons there. Then, again, the island seems to have been elevated. I wonder much whether it stands out in the line of any oceanic current, which does not so forcibly strike the main island? But why, oh, why should so many monocotyledons have come there? or why should they have survived there more than on the main island, if once connected? So, again, I cannot conceive that four snakes should have become extinct in Mauritius and survived on Round Island. For a moment I thought that Mauritius might be the newer island, but the enormous degradation which the outer ring of rocks has undergone flatly contradicts this, and the marine remains on the summit of Round Island indicate the island to be comparatively new — unless, indeed, they are fossil and extinct marine remains. Do tell me what you think. There never was such an enigma. I rather lean to separate immigration, with, of course, subsequent modification; some forms, of course, also coming from Mauritius. Speaking of Mauritius reminds me that I was so much pleased the day before yesterday by reading a review of a book on the geology of St. Helena, by an officer who knew nothing of my hurried observations, but confirms nearly all that I have said on the general structure of the island, and on its marvellous denudation. The geology of that island was like a novel.
LETTER 387. TO A. BLYTT. Down, March 28th, 1876.
(387/1. The following refers to Blytt's "Essay on the Immigration of the Norwegian Flora during Alternating Rainy and Dry Periods," Christiania, 1876.)
I thank you sincerely for your kindness in having sent me your work on the "Immigration of the Norwegian Flora," which has interested me in the highest degree. Your view, supported as it is by various facts, appears to me the most important contribution towards understanding the present distribution of plants, which has appeared since Forbes' essay on the effects of the Glacial Period.
LETTER 388. TO AUG. FOREL. Down, June 19th, 1876.
I hope you will allow me to suggest an observation, should any opportunity occur, on a point which has interested me for many years — viz., how do the coleoptera which inhabit the nests of ants colonise a new nest? Mr. Wallace, in reference to the presence of such coleoptera in Madeira, suggests that their ova may be attached to the winged female ants, and that these are occasionally blown across the ocean to the island. It would be very interesting to discover whether the ova are adhesive, and whether the female coleoptera are guided by instinct to attach them to the female ants (388/1. Dr. Sharp is good enough to tell us that he is not aware of any such adaptation. Broadly speaking, the distribution of the nest-inhabiting beetles is due to co-migration with the ants, though in some cases the ants transport the beetles. Sitaris and Meloe are beetles which live "at the expense of bees of the genus Anthophora." The eggs are laid not in but near the bees' nest; in the early stage the larva is active and has the instinct to seize any hairy object near it, and in this way they are carried by the Anthophora to the nest. Dr. Sharp states that no such preliminary stage is known in the ant's-nest beetles. For an account of Sitaris and Meloe, see Sharp's "Insects," II., page 272.); or whether the larvae pass through an early stage, as with Sitaris or Meloe, or cling to the bodies of the females. This note obviously requires no answer. I trust that you continue your most interesting investigations on ants.
(PLATE: MR. A.R. WALLACE, 1878. From a photograph by Maull & Fox.)
LETTER 389. TO A.R. WALLACE.
(389/1. Published in "Life and Letters," III., page 230.)
(389/2. The following five letters refer to Mr. Wallace's "Geographical Distribution of Animals," 1876.)
{Hopedene} (389/3. Mr. Hensleigh Wedgwood's house in Surrey.), June 5th, 1876.
I must have the pleasure of expressing to you my unbounded admiration of your book (389/4. "Geographical Distribution," 1876.), though I have read only to page 184 — my object having been to do as little as possible while resting. I feel sure that you have laid a broad and safe foundation for all future work on Distribution. How interesting it will be to see hereafter plants treated in strict relation to your views; and then all insects, pulmonate molluscs and fresh-water fishes, in greater detail than I suppose you have given to these lower animals. The point which has interested me most, but I do not say the most valuable point, is your protest against sinking imaginary continents in a quite reckless manner, as was stated by Forbes, followed, alas, by Hooker, and caricatured by Wollaston and {Andrew} Murray! By the way, the main impression that the latter author has left on my mind is his utter want of all scientific judgment. I have lifted up my voice against the above view with no avail, but I have no doubt that you will succeed, owing to your new arguments and the coloured chart. Of a special value, as it seems to me, is the conclusion that we must determine the areas, chiefly by the nature of the mammals. When I worked many years ago on this subject, I doubted much whether the now-called Palaearctic and Nearctic regions ought to be separated; and I determined if I made another region that it should be Madagascar. I have, therefore, been able to appreciate your evidence on these points. What progress Palaeontology has made during the last twenty years! but if it advances at the same rate in the future, our views on the migration and birthplace of the various groups will, I fear, be greatly altered. I cannot feel quite easy about the Glacial period, and the extinction of large mammals, but I must hope that you are right. I think you