The Malay Archipelago, Volume 1. Alfred Russel Wallace
changes did occur in the organic world.
Taking, therefore, these two series of changes for granted, most of the present peculiarities and anomalies in the distribution of species may be directly traced to them. In our own islands, with a very few trifling exceptions, every quadruped, bird, reptile, insect, and plant, is found also on the adjacent continent. In the small islands of Sardinia and Corsica, there are some quadrupeds and insects, and many plants, quite peculiar. In Ceylon, more closely connected to India than Britain is to Europe, many animals and plants are different from those found in India, and peculiar to the island. In the Galapagos Islands, almost every indigenous living thing is peculiar to them, though closely resembling other kinds found in the nearest parts of the American continent.
Most naturalists now admit that these facts can only be explained by the greater or less lapse of time since the islands were upraised from beneath the ocean, or were separated from the nearest land; and this will be generally (though not always) indicated by the depth of the intervening sea. The enormous thickness of many marine deposits through wide areas shows that subsidence has often continued (with intermitting periods of repose) during epochs of immense duration. The depth of sea produced by such subsidence will therefore generally be a measure of time; and in like manner, the change which organic forms have undergone is a measure of time. When we make proper allowance for the continued introduction of new animals and plants from surrounding countries by those natural means of dispersal which have been so well explained by Sir Charles Lyell and Mr. Darwin, it is remarkable how closely these two measures correspond. Britain is separated from the continent by a very shallow sea, and only in a very few cases have our animals or plants begun to show a difference from the corresponding continental species. Corsica and Sardinia, divided from Italy by a much deeper sea, present a much greater difference in their organic forms. Cuba, separated from Yucatan by a wider and deeper strait, differs more markedly, so that most of its productions are of distinct and peculiar species; while Madagascar, divided from Africa by a deep channel three hundred miles wide, possesses so many peculiar features as to indicate separation at a very remote antiquity, or even to render it doubtful whether the two countries have ever been absolutely united.
Returning now to the Malay Archipelago, we find that all the wide expanse of sea which divides Java, Sumatra, and Borneo from each other, and from Malacca and Siam, is so shallow that ships can anchor in any part of it, since it rarely exceeds forty fathoms in depth; and if we go as far as the line of a hundred fathoms, we shall include the Philippine Islands and Bali, east of Java. If, therefore, these islands have been separated from each other and the continent by subsidence of the intervening tracts of land, we should conclude that the separation has been comparatively recent, since the depth to which the land has subsided is so small. It is also to be remarked that the great chain of active volcanoes in Sumatra and Java furnishes us with a sufficient cause for such subsidence, since the enormous masses of matter they have thrown out would take away the foundations of the surrounding district; and this may be the true explanation of the often-noticed fact that volcanoes and volcanic chains are always near the sea. The subsidence they produce around them will, in time, make a sea, if one does not already exist.
But, it is when we examine the zoology of these countries that we find what we most require—evidence of a very striking character that these great islands must have once formed a part of the continent, and could only have been separated at a very recent geological epoch. The elephant and tapir of Sumatra and Borneo, the rhinoceros of Sumatra and the allied species of Java, the wild cattle of Borneo and the kind long supposed to be peculiar to Java, are now all known to inhabit some part or other of Southern Asia. None of these large animals could possibly have passed over the arms of the sea which now separate these countries, and their presence plainly indicates that a land communication must have existed since the origin of the species. Among the smaller mammals, a considerable portion are common to each island and the continent; but the vast physical changes that must have occurred during the breaking up and subsidence of such extensive regions have led to the extinction of some in one or more of the islands, and in some cases there seems also to have been time for a change of species to have taken place. Birds and insects illustrate the same view, for every family and almost every genus of these groups found in any of the islands occurs also on the Asiatic continent, and in a great number of cases the species are exactly identical. Birds offer us one of the best means of determining the law of distribution; for though at first sight it would appear that the watery boundaries which keep out the land quadrupeds could be easily passed over by birds, yet practically it is not so; for if we leave out the aquatic tribes which are pre-eminently wanderers, it is found that the others (and especially the Passeres, or true perching-birds, which form the vast majority) are generally as strictly limited by straits and arms of the sea as are quadrupeds themselves. As an instance, among the islands of which I am now speaking, it is a remarkable fact that Java possesses numerous birds which never pass over to Sumatra, though they are separated by a strait only fifteen miles wide, and with islands in mid-channel. Java, in fact, possesses more birds and insects peculiar to itself than either Sumatra or Borneo, and this would indicate that it was earliest separated from the continent; next in organic individuality is Borneo, while Sumatra is so nearly identical in all its animal forms with the peninsula of Malacca, that we may safely conclude it to have been the most recently dismembered island.
The general result therefore, at which we arrive, is that the great islands of Java, Sumatra, and Borneo resemble in their natural productions the adjacent parts of the continent, almost as much as such widely-separated districts could be expected to do even if they still formed a part of Asia; and this close resemblance, joined with the fact of the wide extent of sea which separates them being so uniformly and remarkably shallow, and lastly, the existence of the extensive range of volcanoes in Sumatra and Java, which have poured out vast quantities of subterranean matter and have built up extensive plateaux and lofty mountain ranges, thus furnishing a vera causa for a parallel line of subsidence—all lead irresistibly to the conclusion that at a very recent geological epoch, the continent of Asia extended far beyond its present limits in a south-easterly direction, including the islands of Java, Sumatra, and Borneo, and probably reaching as far as the present 100-fathom line of soundings.
The Philippine Islands agree in many respects with Asia and the other islands, but present some anomalies, which seem to indicate that they were separated at an earlier period, and have since been subject to many revolutions in their physical geography.
Turning our attention now to the remaining portion of the Archipelago, we shall find that all the islands from Celebes and Lombock eastward exhibit almost as close a resemblance to Australia and New Guinea as the Western Islands do to Asia. It is well known that the natural productions of Australia differ from those of Asia more than those of any of the four ancient quarters of the world differ from each other. Australia, in fact, stands alone: it possesses no apes or monkeys, no cats or tigers, wolves, bears, or hyenas; no deer or antelopes, sheep or oxen; no elephant, horse, squirrel, or rabbit; none, in short, of those familiar types of quadruped which are met with in every other part of the world. Instead of these, it has Marsupials only: kangaroos and opossums; wombats and the duckbilled Platypus. In birds it is almost as peculiar. It has no woodpeckers and no pheasants—families which exist in every other part of the world; but instead of them it has the mound-making brush-turkeys, the honeysuckers, the cockatoos, and the brush-tongued lories, which are found nowhere else upon the globe. All these striking peculiarities are found also in those islands which form the Austro-Malayan division of the Archipelago.
The great contrast between the two divisions of the Archipelago is nowhere so abruptly exhibited as on passing from the island of Bali to that of Lombock, where the two regions are in closest proximity. In Bali we have barbets, fruit-thrushes, and woodpeckers; on passing over to Lombock these are seen no more, but we have abundance of cockatoos, honeysuckers, and brush-turkeys, which are equally unknown in Bali, or any island further west. [I was informed, however, that there were a few cockatoos at one spot on the west of Bali, showing that the intermingling of the productions of these islands is now going on.] The strait is here fifteen miles wide, so that we may pass in two hours from one great division of the earth to another, differing as essentially in their animal life as Europe does from America. If we travel from Java or Borneo to Celebes or the Moluccas, the difference is still more striking. In the first, the forests abound in monkeys of many kinds, wild cats, deer, civets, and