The Voyage of the Beagle. Чарльз Дарвин
alongside each other, it is not easy to conceive ranks more disproportionate in size. After the above facts, we are compelled to conclude, against anterior probability, 58 that among the mammalia there exists no close relation between the bulk of the species, and the quantity of the vegetation, in the countries which they inhabit.
With regard to the number of large quadrupeds, there certainly exists no quarter of the globe which will bear comparison with Southern Africa. After the different statements which have been given, the extremely desert character of that region will not be disputed. In the European division of the world, we must look back to the tertiary epochs, to find a condition of things among the mammalia, resembling that now existing at the Cape of Good Hope. Those tertiary epochs, which we are apt to consider as abounding to an astonishing degree with large animals, because we find the remains of many ages accumulated at certain spots, could hardly boast of more large quadrupeds than Southern Africa does at present. If we speculate on the condition of the vegetation during these epochs we are at least bound so far to consider existing analogies, as not to urge as absolutely necessary a luxuriant vegetation, when we see a state of things so totally different at the Cape of Good Hope.
We know 59 that the extreme regions of North America, many degrees beyond the limit where the ground at the depth of a few feet remains perpetually congealed, are covered by forests of large and tall trees. In a like manner, in Siberia, we have woods of birch, fir, aspen, and larch, growing in a latitude 510 (64 degs.) where the mean temperature of the air falls below the freezing point, and where the earth is so completely frozen, that the carcass of an animal embedded in it is perfectly preserved. With these facts we must grant, as far as quantity alone of vegetation is concerned, that the great quadrupeds of the later tertiary epochs might, in most parts of Northern Europe and Asia, have lived on the spots where their remains are now found. I do not here speak of the kind of vegetation necessary for their support; because, as there is evidence of physical changes, and as the animals have become extinct, so may we suppose that the species of plants have likewise been changed.
These remarks, I may be permitted to add, directly bear on the case of the Siberian animals preserved in ice. The firm conviction of the necessity of a vegetation possessing a character of tropical luxuriance, to support such large animals, and the impossibility of reconciling this with the proximity of perpetual congelation, was one chief cause of the several theories of sudden revolutions of climate, and of overwhelming catastrophes, which were invented to account for their entombment. I am far from supposing that the climate has not changed since the period when those animals lived, which now lie buried in the ice. At present I only wish to show, that as far as quantity of food alone is concerned, the ancient rhinoceroses might have roamed over the steppes of central Siberia (the northern parts probably being under water) even in their present condition, as well as the living rhinoceroses and elephants over the Karros of Southern Africa.
I will now give an account of the habits of some of the more interesting birds which are common on the wild plains of Northern Patagonia: and first for the largest, or South American ostrich. The ordinary habits of the ostrich are familiar to every one. They live on vegetable matter, such as roots and grass; but at Bahia Blanca I have repeatedly seen three or four come down at low water to the extensive mud-banks which are then dry, for the sake, as the Gauchos say, of feeding on small fish. Although the ostrich in its habits is so shy, wary, and solitary, and although so fleet in its pace, it is caught without much difficulty by the Indian or Gaucho armed with the bolas. When several horsemen appear in a semicircle, it becomes confounded, and does not know which way to escape. They generally prefer running against the wind; yet at the first start they expand their wings, and like a vessel make all sail. On one fine hot day I saw several ostriches enter a bed of tall rushes, where they squatted concealed, till quite closely approached. It is not generally known that ostriches readily take to the water. Mr. King informs me that at the Bay of San Blas, and at Port Valdes in Patagonia, he saw these birds swimming several times from island to island. They ran into the water both when driven down to a point, and likewise of their own accord when not frightened: the distance crossed was about two hundred yards. When swimming, very little of their bodies appear above water; their necks are extended a little forward, and their progress is slow. On two occasions I saw some ostriches swimming across the Santa Cruz river, where its course was about four hundred yards wide, and the stream rapid. Captain Sturt, 511 when descending the Murrumbidgee, in Australia, saw two emus in the act of swimming.
The inhabitants of the country readily distinguish, even at a distance, the cock bird from the hen. The former is larger and darker-coloured, 512 and has a bigger head. The ostrich, I believe the cock, emits a singular, deep-toned, hissing note: when first I heard it, standing in the midst of some sand-hillocks, I thought it was made by some wild beast, for it is a sound that one cannot tell whence it comes, or from how far distant. When we were at Bahia Blanca in the months of September and October, the eggs, in extraordinary numbers, were found all over the country. They lie either scattered and single, in which case they are never hatched, and are called by the Spaniards huachos; or they are collected together into a shallow excavation, which forms the nest. Out of the four nests which I saw, three contained twenty-two eggs each, and the fourth twenty-seven. In one day's hunting on horseback sixty-four eggs were found; forty-four of these were in two nests, and the remaining twenty, scattered huachos. The Gauchos unanimously affirm, and there is no reason to doubt their statement, that the male bird alone hatches the eggs, and for some time afterwards accompanies the young. The cock when on the nest lies very close; I have myself almost ridden over one. It is asserted that at such times they are occasionally fierce, and even dangerous, and that they have been known to attack a man on horseback, trying to kick and leap on him. My informer pointed out to me an old man, whom he had seen much terrified by one chasing him. I observe in Burchell's travels in South Africa, that he remarks, "Having killed a male ostrich, and the feathers being dirty, it was said by the Hottentots to be a nest bird." I understand that the male emu in the Zoological Gardens takes charge of the nest: this habit, therefore, is common to the family.
The Gauchos unanimously affirm that several females lay in one nest. I have been positively told that four or five hen birds have been watched to go in the middle of the day, one after the other, to the same nest. I may add, also, that it is believed in Africa, that two or more females lay in one nest. 513 Although this habit at first appears very strange, I think the cause may be explained in a simple manner. The number of eggs in the nest varies from twenty to forty, and even to fifty; and according to Azara, some times to seventy or eighty. Now, although it is most probable, from the number of eggs found in one district being so extraordinarily great in proportion to the parent birds, and likewise from the state of the ovarium of the hen, that she may in the course of the season lay a large number, yet the time required must be very long. Azara states, 514 that a female in a state of domestication laid seventeen eggs, each at the interval of three days one from another. If the hen was obliged to hatch her own eggs, before the last was laid the first probably would be addled; but if each laid a few eggs at successive periods, in different nests, and several hens, as is stated to be the case, combined together, then the eggs in one collection would be nearly of the same age. If the number of eggs in one of these nests is, as I believe, not greater on an average than the number laid by one female in the season, then there must be as many nests as females, and each cock bird will have its fair share of the labour of incubation; and that during a period when the females probably could not sit, from not having finished laying. 515 I have before mentioned the great numbers of huachos, or deserted eggs; so that in one day's hunting twenty were found in this state. It appears odd that so many should be wasted. Does it not arise from the difficulty of several females associating together, and finding a male ready to undertake the office of incubation? It is evident that there must at first be some degree of association between at least two females; otherwise the eggs would remain scattered over the wide plain, at distances far too great to allow of the male collecting them into one nest: some authors have believed that the scattered eggs were deposited for the young birds to feed on. This can hardly be the case in America, because the huachos, although often found addled and putrid, are generally whole.
When at the Rio Negro in Northern Patagonia, I repeatedly heard the Gauchos talking of a very rare bird which they called Avestruz Petise. They described it as being less than the common ostrich (which is there abundant), but with