The Physical Basis of Mind. George Henry Lewes
“emboîtement” theory. In spite of obvious objections it gained scientific acceptance, because physiologists could not bring themselves to believe that so marvellous a structure as that of a human organism arose by a series of successive modifications, or because they could not comprehend how it was built up, part by part, into forms so closely resembling the parent-forms. That many and plausible reasons pleaded in favor of this opinion is evident in the fact that illustrious men like Haller, Bonnet, Vallisneri, Swammerdamm, Réaumur, and Cuvier, were its advocates; and if there is not a sigle physiologist of our day who accepts it, or who finds any peculiar difficulty in following the demonstrations of embryologists, how from the common starting-point of a self-multiplying epithelial cell parts so diverse as hairs, nails, hoofs, scales, feathers, crystalline lens, and secreting glands may be evolved, or how from the homogeneous germinal membrane the complex organism will arise, there are very few among the scorners of the dead hypothesis who seem capable of generalizing the principles which have destroyed it, or can conceive that the laws of Evolution apply as rigorously to the animal and vegetable kingdoms as to the individual organisms. The illustrious names of those who advocated the preformation hypothesis may serve to check our servile submission to the authorities so loudly proclaimed as advocates of the fixity of species. The more because the two doctrines have a common parentage. The one falls with the other, and no array of authorities can arrest the fall. That the manifold differentiations noticeable in a complex organism should have been evolved from a membrane wholly destitute of differences is a marvel, but a marvel which Science has made intelligible. Yet the majority of those to whom this has been made intelligible still find an impossibility in admitting that the manifold forms of plant and animal were successively evolved from equally simple origins. They relinquish the hypothesis of preformation in the one case, and cling to it in the other. Evolution, demonstrable in the individual history, seems preposterous in the history of the class. And thus is presented the instructive spectacle of philosophers laughing at the absurdities of “preformation,” and yet exerting all their logic and rhetoric in defence of “creative fiats”—which is simply the preformation hypothesis “writ large.”
101. It would not be difficult to show that the doctrine of Epigenesis, with which Wolff forever displaced the doctrine of Preformation, leads by an inevitable logic to the doctrine of universal Evolution; and that we can no more understand the appearance of a new organism which is not the modification of some already existing organism, than we can understand the sudden appearance of a new organ which is not the modification of some existing structure. In the one case as in the other we may disguise the process under such terms as creative fiat and preformation; but these terms are no explanations; they re-state the results, they do not describe the process; whereas Epigenesis describes the process as it passes under the eye of science.
102. If any reader of these pages who, from theological or zoölogical suspicion of the Development Hypothesis, clings to the hypothesis of a creative Plan which once for all arranged the organic world in Types that could not change, will ask what rational interpretation can be given to the succession of phases each embryo is forced to pass through, it may help to give him pause. He will observe that none of these phases have any adaptation to the future state of the animal, but are in positive contradiction to it, or are simply purposeless; whereas all show stamped on them the unmistakable characters of ancestral adaptations and the progressions of Organic Evolution. What does the fact imply? There is not a single known example of a complex organism which is not developed out of simpler forms. Before it can attain the complex structure which distinguishes it, there must be an evolution of forms similar to those which distinguish the structures of organisms lower in the series. On the hypothesis of a Plan which prearranged the organic world, nothing could be more unworthy of a supreme intelligence than this inability to construct an organism at once, without previously making several tentative efforts, undoing to-day what was so carefully done yesterday, and repeating for centuries the same tentatives, and the same corrections, in the same succession. Do not let us blink this consideration. There is a traditional phrase much in vogue among the anthropomorphists, which arose naturally enough from the tendency to take human methods as an explanation of the divine—a phrase which becomes a sort of argument—“The Great Architect.” But if we are to admit the human point of view, a glance at the facts of embryology must produce very uncomfortable reflections. For what should we say to an architect who was unable, or being able was obstinately unwilling, to erect a palace except by first using his materials in the shape of a hut, then pulling it down and rebuilding them as a cottage, then adding story to story and room to room, not with any reference to the ultimate purposes of the palace, but wholly with reference to the way in which houses were constructed in ancient times? What should we say to the architect who could not form a museum out of bricks and mortar, but was forced to begin as if going to build a mansion: and after proceeding some way in this direction, altered his plan into a palace, and that again into a museum? Yet this is the sort of succession on which organisms are constructed. The fact has long been familiar; how has it been reconciled with Infinite Wisdom? Let the following passage answer for a thousand:—“The embryo is nothing like the miniature of the adult. For a long while the body in its entirety and its details presents the strangest of spectacles. Day by day and hour by hour the aspect of the scene changes, and this instability is exhibited by the most essential parts no less than by the accessory parts. One would say that Nature feels her way, and only reaches the goal after many times missing the path—on dirait que la nature tâtonne et ne conduit son œuvre à bon fin qu’après s’être souvent trompée.”45 Writers have no compunction in speaking of Nature feeling her way and blundering; but if in lieu of Nature, which may mean anything, the Great Architect be substituted, it is probable that the repugnance to using such language of evasion may cause men to revise their conceptions altogether; they dare not attribute ignorance and incompetence to the Creator.
103. Obviously the architectural hypothesis is incompetent to explain the phenomena of organic development. Evolution is the universal process; not creation of a direct kind. Von Baer, who very properly corrected the exaggerations which had been put forth respecting the identity of the embryonic forms with adult forms lower in the scale, who showed that the mammalian embryo never was a bird, a reptile, or a fish, nevertheless emphasized the fact that the mammalian embryo passes through all the lower typical forms; so much so that, except by their size, it is impossible to distinguish the embryos of mammal, bird, lizard, or snake. “In my collection,” he says, “there are two little embryos which I have omitted to label, so that I am now quite incompetent to say to what class they belong. They may be lizards, they may be small birds, or very young mammals; so complete is the similarity in the mode of formation of the head and trunk. The extremities have not yet made their appearance. But even if they existed in the earliest stage we should learn nothing from them, for the feet of lizards, mammals, and the wings of birds, all arise from the same common form.” He sums up with his formula: “The special type is always evolved from a more general type.”46
Such reminiscences of earlier forms are intelligible on the supposition that originally the later form was a modification of the earlier form, and that this modification is repeated; or on the supposition that there was a similarity in the organic conditions, which similarity ceased at the point where the new form emerged. But on no hypothesis of creative Plan are they intelligible. They are useless structures, failing even to subserve a temporary purpose. Sometimes, as Mr. Darwin remarks, a trace of the embryonic resemblance lasts till a late age: “Thus birds of the same genus, and of closely allied genera, often resemble each other in their first and second plumage: as we see in the spotted feathers in the thrush group. In the cat tribe most of the species are striped and spotted in lines; and stripes or spots can plainly be distinguished in the whelp of the lion and the puma. We occasionally, though rarely, see something of this kind in plants. … The points of structure in which the embryos of widely different animals of the same class resemble each other often have no direct relation to their conditions of existence. We cannot, for instance, suppose that in the embryos of the vertebrata the peculiar loop-like courses of the arteries near the bronchial slits are related to similar conditions in the young mammal which is nourished in the womb of its mother, in the egg of a bird which is hatched in a nest, and in the spawn of a frog under water.”
104. It would be easy to multiply examples, but I will content myself with three. The tadpole of the Salamander